Calobata guttata, (Walker)

Ferro, Gustavo Borges & Marshall, Stephen A., 2018, A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae), Zootaxa 4429 (3), pp. 401-458: 407-409

publication ID

https://doi.org/10.11646/zootaxa.4429.3.1

publication LSID

lsid:zoobank.org:pub:004C0DCB-9B63-4A44-A1BB-2D98ADFB1CEE

persistent identifier

http://treatment.plazi.org/id/0A6A87F2-FFC5-802C-FF10-FB49FA61FE6D

treatment provided by

Plazi

scientific name

Calobata guttata
status

 

1 Wing dark with three round hyaline spots on apical third ( Fig. 40 View Figure ); discal band sometimes dark with up to four hyaline spots ( Fig. 84 View Figure )............................................................................................ 2

- Wing apex either entirely dark ( Fig. 86 View Figure ) or clear ( Fig. 103 View Figure ) or clear with a narrow preapical pigmentation ( Fig. 62 View Figure ) or three preapical black spots ( Figs. 35 View Figure , 43 View Figure , 74 View Figure ); discal band entirely dark................................................ 3

2 Paracephalon swollen and conical ( Fig. 40 View Figure ). Vertex flattened. Wing with discal band black, without hyaline spots............................ C. guttata (Walker)  . Brazil, Colombia, French Guiana, Guiana, Surinam, Panama, Trinidad, Venezuela

- Paracephalon without swelling ( Fig. 82 View Figure ). Vertex rounded. Wing with discal band dissected by four hyaline rounded spots................................................................................. C. secunda Hennig.  Ecuador

3 Wing apex entirely clear ( Fig. 103 View Figure )..................................................... C. vitrata  n. sp. Ecuador

- Wing apex not as above, either entirely dark ( Fig. 86 View Figure ) or clear with a narrow preapical pigmentation ( Fig. 62 View Figure ) or three preapical black spots ( Figs. 35 View Figure , 43 View Figure , 74 View Figure )............................................................................ 4

4 Clypeus long, length more than half of width in frontal view ( Fig. 90 View Figure ). Fore tarsomeres 1–3 white (tarsomere one with brown pubescence basally), 4–5 covered with brown pubescence ( Figs. 28 View Figure , 92 View Figure ).......................................... 5

- Clypeus short, length less than half of width in frontal view. Fore tarsus not as above (either tarsomere one dark brown with gold/whitish ventral pubescence basally ( Fig. 9 View Figure ) OR fore tarsomere one dark brown and 2–5 white ( Fig. 30 View Figure ) OR fore tarsomere one white and 2–5 dark brown ( Fig. 24 View Figure )).................................................................. 6

5 Thorax brown. Paired spermathecal duct splitting into two convoluted stems marked by several long finger-like process ( Fig. 91 View Figure ).............................................................................. C. spinosa  n. sp. Brazil

- Thorax black. Paired spermathecal duct splitting into two convoluted and broadened stems, without finger-like process ( Fig. 29 View Figure )......................................................................... C. camptomera (Bigot)  . Brazil

6 Three frontal setae ( Fig. 4 View Figure ). Fore tarsomere one dark brown, tarsomeres 2–5 white.................................. 7

- Two frontal setae ( Fig. 14 View Figure ). Fore tarsomeres 2–5 dark brown................................................... 9

7 Scutellum upturned and peaked, slightly elongated at apex ( Fig. 4 View Figure )............ C. acuminata  n. sp. Colombia, Costa Rica

- Scutellum upturned and broadly triangular ( Figs. 30 View Figure , 86 View Figure )..................................................... 8

8 Paracephalon swollen, swelling rounded ( Fig. 30 View Figure ). Abdominal pleural membrane dark brown, sometimes with rusty red regions. Yellowish species...................... C. cinnameus (Hennig)  . Bolivia, Brazil, Colombia, French Guiana, Peru

- Paracephalon smooth, without swelling. Abdominal pleural membrane black. Blackish species.............................................................................. C. setosa (Hennig)  . Bolivia, Brazil, Colombia, Ecuador, Peru

9 Mid and hind tibiae flattened, sulcus on outer face strongly pronounced and curved ( Figs. 18 View Figure , 48 View Figure , 55 View Figure , 97 View Figure ). Blackish species...................................................................................................... 10

- Mid and hind tibiae slightly flattened, sulcus on outer face weak. Colour variable.................................. 13

10 Inner vertical setae insertions slightly protuberant ( Fig. 16 View Figure , 46 View Figure ). Wing apex clear with a dark and narrow preapical pigmentation................................................................................................ 11

- Inner vertical setae insertions flat. Wing apex darkened....................................................... 12

11 Paracephalon swollen ( Fig. 16 View Figure ). Paired spermathecae flat, disc-shaped...... C. aspera  n. sp. Bolivia, Ecuador, Guiana, Peru

- Paracephalon without swelling ( Fig. 46 View Figure ). Paired spermathecae spherical................................................................................................ C. lobulosa  n. sp. Bolivia, Brazil, Colombia, Ecuador, Peru

12 Epicephalon black ( Fig. 55 View Figure ). Ejaculatory apodeme large, blade wider than sperm pump.......................................... C. nigra Schiner.  Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Honduras, Mexico, Peru, Trinidad, Venezuela

- Epicephalon velvet brown ( Fig. 97 View Figure ). Ejaculatory apodeme small, sperm pump wider than blade..... C. velutinosa  n. sp. Peru

13 Mid and hind tibia bicoloured, basal half dark brown and apical half yellowish ( Fig. 59 View Figure ). Crossvein r-m clear, without dark spot........................................................................ C. peruana (Hennig)  . Bolivia, Peru

- Mid and hind tibia uniformly black, brown or yellowish. Crossvein r-m with small dark spot......................... 14

14 Paracephalon swollen ( Figs. 10 View Figure , 24 View Figure , 33 View Figure , 72 View Figure ) or slightly swollen ( Fig. 38 View Figure ), extending conically beyond the vertex........ 15

- Paracephalon without swelling, not extending conically beyond the vertex...................................... 19

15 Limits between paracephalon, epicephalon and vertex not distinguishable ( Fig. 38 View Figure ). Ocellar triangle displaced anteriorly (closer to anterior frons margin), vertex occupying more than half the length of the head.... C. erugata  n. sp. Ecuador, Peru

- Limits between paracephalon, epicephalon and vertex clearly distinguishable. Ocellar triangle centralized, vertex occupying no more than half the length of the head................................................................... 16

16 Inner vertical setae inserted in strong tubercles ( Fig. 33 View Figure ). A1+CuA2 less than half as long as CuA2............................................................................... C. conifera Hendel.  Brazil, Colombia, Guiana, Paraguay

- Inner vertical seta insertion flat, without tubercle. A1+CuA2 as long as CuA2..................................... 17

17 Female cervical sclerite with a small nipple-like swelling in posterior half. Wing apex clear with three preapical black spots ( Fig. 72 View Figure ). Black species...................................... C. podagrica Rondani.  Bolivia, Brazil, Ecuador, Peru

- Female cervical sclerite smooth, without nipple-like swelling in posterior half. Wing apex clear with a dark and narrow preapical pigmentation. Yellowish to brown species, sometimes slightly rusty red....................................... 18

18 Swelling on paracephalon pointing dorsally ( Fig. 9 View Figure ). Fore tarsomere one dark brown with gold/whitish ventral pubescence basally. Yellowish species............................................................ C. angularis  n. sp. Peru

- Swelling on paracephalon posteriorly pointed ( Figs. 24, 26 View Figure ). Fore tarsomere one white, fore tarsomeres 2–5 dark brown. Brown, sometimes slightly rusty red species................... C. bulla  n. sp. Bolivia, Brazil, Colombia, Ecuador, Peru

19 Vertex flattened, head depressed posteriorly ( Figs. 7 View Figure , 68 View Figure , 74 View Figure , 93 View Figure )................................................ 20

- Vertex rounded, head globose ( Figs. 14 View Figure , 43 View Figure , 50 View Figure , 53 View Figure , 64 View Figure , 78 View Figure , 80 View Figure , 99 View Figure )............................................. 24

20 Mid femur white in basal third. Black/dark brown species.................................................... 21

- Mid femur uniformly brown in basal third. Brown/yellowish species........................................... 22

21 Fore tarsomere one white, fore tarsomeres 2–5 dark brown. Wing apex clear with three preapical black spots. Phallic bulb subequal in length and width, almost spherical ( Fig. 96 View Figure ).......... C. umbra  n. sp. French Guiana, Tobago, Trinidad, Venezuela

- Fore tarsomere one dark brown with gold/whitish ventral pubescence basally. Wing apex clear with a dark and narrow preapical pigmentation. Phallic bulb cylindrical, about three times as long as wide ( Fig. 23 View Figure ).......................................................... C. brevipennis (Walker)  . Bolivia, Brazil, Colombia, Ecuador, French Guiana, Peru, Venezuela

22 Epicephalon smooth, without wrinkles posterior to ocellar triangle ( Fig. 74 View Figure ). Fore tarsomere one white, fore tarsomeres 2–5 dark brown. Wing apex clear with three preapical black spots....................... C. punctipennis (Macquart)  . Brazil

- Epicephalon wrinkled posterior to ocellar triangle ( Figs. 7 View Figure , 68 View Figure ). Fore tarsomere one dark brown with gold/whitish ventral pubescence basally. Wing apex clear with a dark and narrow preapical pigmentation............................... 23

23 Yellowish species. Paracephalon and epicephalon yellow ( Fig. 68 View Figure ). Paired spermathecae cylindrical and thin ( Fig. 67 View Figure ). Phallic bulb cylindrical, about three times as long as wide ( Fig. 69 View Figure ).................................................................................................. C. planivertex  n. sp. Brazil, Colombia, Ecuador, French Guiana, Peru

- Brownish species, head and abdomen usually rusty red. Paracephalon and epicephalon darkened ( Figs. 5, 7 View Figure ). Paired spermathecae sac-like, tapered basally and apically swollen ( Fig. 6 View Figure ). Phallic bulb subequal in length and width, almost spherical ( Fig. 8 View Figure )............................................................................. C. aeruginosa  n. sp. Ecuador

24 Inner vertical setae insertions with strong tubercles ( Fig. 50 View Figure )................ C. modesta (Cresson)  . Colombia, Guatemala

- Inner vertical setae insertions flat, without tubercles........................................................ 25

25 Fore tarsomere one white, fore tarsomeres 2–5 dark brown................................................... 26

- Fore tarsomere one dark brown with gold/whitish ventral pubescence basally..................................... 27

26 Vertex brownish anterior to postocellar setae. Mid femur uniformly brown in basal third. Thorax brown....................................................................................... C. rotunda  n. sp. Colombia, Costa Rica

- Vertex with a rounded black spot anterior to postocellar setae ( Fig. 53 View Figure ). Mid femur white in basal third. Thorax dark brown with blue metallic shine...................................................... C. nicaraguensis  n. sp. Nicaragua

27 Female cervical sclerite slightly swollen posteriorly. Mid and hind femur white basally, apical third with an angled whitish yellow ring ( Figs. 62, 64 View Figure ).......................................... C. picina (Cresson)  . Costa Rica, Ecuador, Panama

- Female cervical sclerite flat, without posterior swelling. Mid and hind femur with basal and apical third brown or yellow. 28

28 R4+5 and M1+2 ending separately in wing margin. Paired spermathecae cylindrical and strongly curved in middle ( Fig. 100 View Figure )............................................................................. C. venezuelensis  n. sp. Venezuela

- R4+5 and M1+2 meeting at wing margin. Paired spermathecae spherical or sac-like (tapered basally and distally swollen).... 29

29 Epicephalon smooth and shiny, not depressed anterior to postocellar setae ( Fig. 14 View Figure ). Yellowish species, sometimes with abdomen darkened at least on T4–T6.......................................................................................... C. arthritica (Wiedemann)  . Belize, Colombia, Costa Rica, Ecuador, Honduras, Mexico, Panama, Venezuela

- Epicephalon somewhat wrinkled and depressed before postocellar setae ( Figs. 43 View Figure , 80 View Figure ). Brownish/ferruginous species with darkened regions on mesonotum and katepisternum........................................................ 30

30 Wing apex clear with three preapical black spots. Paired spermathecal duct stems with a preapical rounded swelling ( Fig. 44 View Figure ).................................... C. harenosa (Cresson)  . Colombia, Costa Rica, Ecuador, Guiana, Mexico, Panama

- Wing apex clear with a dark and narrow preapical band. Paired spermathecal duct stems very narrow in basal third, then expanded and convoluted ( Fig. 81 View Figure )............................................... C. schildi (Cresson)  . Costa Rica

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Micropezidae

Genus

Calobata