Cyrtodactylus thuongae , Phung, Trung My, Schingen, Mona Van, Ziegler, Thomas & Nguyen, Truong Quang, 2014

Phung, Trung My, Schingen, Mona Van, Ziegler, Thomas & Nguyen, Truong Quang, 2014, A third new Cyrtodactylus (Squamata: Gekkonidae) from Ba Den Mountain, Tay Ninh Province, southern Vietnam, Zootaxa 3764 (3), pp. 347-363: 348-359

publication ID

http://dx.doi.org/10.11646/zootaxa.3764.3.5

publication LSID

lsid:zoobank.org:pub:677C878F-595E-4639-ADC4-E9F54595BAA5

persistent identifier

http://treatment.plazi.org/id/A12187BD-FF94-8548-EBDB-32EFA116FE21

treatment provided by

Plazi

scientific name

Cyrtodactylus thuongae
status

sp. nov.

Cyrtodactylus thuongae  sp. nov.

Figs. 1View FIGURE 1, 2View FIGURE 2 A, 3

Holotype. IEBR A.2013.23, adult male, collected on 14 May 2013 by T.M. Phung in a rocky cave from Ba Den Mountain, Thanh Tan Ward, Tay Ninh Town, Tay Ninh Province (11 ° 21 ' 52 "N 106 ° 10 ' 27 "E), southern Vietnam, at an elevation of 600 m a.s.l.

Paratypes. IEBR A.2013.24, adult female, IEBR A.2013.25, adult female, VNMNAbout VNMN A.2013.12, adult female, were collected on 16 December 2012; VNMNAbout VNMN A.2013.13, adult male, ZFMKAbout ZFMK 94994–94997, adult females, and ZFMKAbout ZFMK 94998–94999, subadults, were collected on 14 May 2013 by T.M. Phung, the same collection data as the holotype.

Diagnosis. A medium-sized Cyrtodactylus  with a maximum SVL of 77.6 mm, distinguished from all congeners by a combination of the following characters: 1) dorsal pattern consisting of irregular blotches on dorsum and occiput; nuchal loop sometimes interrupted medially; 2) precloacal pore absent or one in males; 3) femoral pore series separated from precloacal pore series, 0–3 on each thigh in males; 4) ventrals in 29–44 longitudinal rows at midbody; 5) lateral skin folds present, without interspersed tubercles; 6) dorsal tubercles in 16–18 irregular rows; 7) enlarged femoral scales and precloacal scales present; 8) postcloacal spurs two or three; 9) subcaudal scales slightly enlarged; and 10) precloacal groove absent.

Description of holotype. Size small (SVL 67.6 mm), tail regenerated (TaL 41.55 mm); supralabials 8 / 9; infralabials 7 / 8; nares bordered by rostral anteriorly, first supralabial laterally and four or five nasals posteriorly; supranasals separated from each other by a pentagonal internasal and two postnasals; upper anterior ciliaries three times larger than posterior ciliaries; dorsal scales of head granular; mental square, as wide as rostral; one pair of enlarged postmentals, longer than wide, in contact with each other posteriorly about a half of their length, bordering mental anteriorly, first two labials laterally, and six gular scales posteriorly; outer pair of gular scales is as large as postmentals; dorsal scales somewhat granular to flattened; dorsal tubercles round, conical, surrounded by 10 granular scales, in 17 irregular longitudinal rows at midbody; ventral scales smooth, in 36 longitudinal rows at midbody; lateral folds present, poorly developed, without interspersed tubercles; upper and lower surface of fore limb without distinct tubercles; dorsal surface of hind limb covered by distinctly developed, round, conical tubercles; fingers and toes lacking distinct webbing; lamellae under fourth finger 15 / 14 and under fourth toe 16 / 17; claws surrounded by a small scale on upper and a large scale on lower sides; series of distinctly enlarged femoral scales separated from enlarged precloacal scales by 15 or 16 small, poreless femoral scales; femoral pores absent; precloacal pore one; enlarged precloacal scales nine, including one pore-bearing scale; precloacal region covered by a patch of approximately 23 enlarged scales behind precloacal pore-bearing scale; precloacal groove absent; postcloacal tubercles 1 / 2, enlarged, on lateral surface of slight hemipenial swelling; hemipenes everted; tail regenerated, dorsum of original tail bearing distinct round tubercles; subcaudals not distinctly enlarged.

Coloration in ethanol. Ground coloration light grey, with dark brown dorsal pattern; dorsum with dark blotches, lighter on the sides; legs, including digits and toes and dorsal surface of tail lighter with dark brown blotches and irregular faint brownish markings between blotches; dorsal surface of head grey with dark irregular blotches on the occiput, snout slightly spotted; venter light cream; ventral surface of tail grey; tubercles on edges of blotches lighter in color; nuchal loop distinct, interrupted and tapering medially. For coloration in life see Fig. 1View FIGURE 1 A.

Variation of paratypes. Two subadults ( ZFMKAbout ZFMK 94998–94999) are uniformly light grey in ground colouration and have distinct brown blotches on dorsum and occiput; limbs and snout are uniformly light grey and their nuchal loops are distinct, complete and tapering medially. The male paratype VNMNAbout VNMN A.2013.13 lacks precloacal and femoral pores. Internasal scales vary from two (in the paratype VNMNAbout VNMN A.2013.13) to three (in the paratypes ZFMKAbout ZFMK 94994, 94995 and 94997). For further variation in scalation see Table 1, and for colour pattern variation see Figure 3View FIGURE 3.

Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Grismer et al. 2008; Rösler & Glaw 2008; Bauer et al. 2009, 2010; Ngo & Grismer 2010; Ngo & Pauwels 2010; Sumontha et al. 2010; Ziegler et al. 2010; David et al. 2011; Iskandar et al. 2011; Luu et al. 2011; Ngo 2011; Ngo & Chan 2011; Schneider et al. 2011; Nazarov et al. 2012; Ngo & Grismer 2012, Ziegler et al. 2013).

Cyrtodactylus thuongae  sp. nov. differs from its Vietnamese congeners by the following characters (see Table 2):

Cyrtodactylus thuongae  sp. nov. has no or one precloacal pore, not in a continuous series with 0–3 femoral pores on each side in males and thus differs from: C. bichnganae Ngo & Grismer 2010  (8–10 PP), C. bidoupimontis Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler 2012  (4–6 PP), C. bugiamapensis Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler 2012  (7 or 8 PP), C. caovansungi Orlov, Nguyen, Nazarov, Ananjeva & Nguyen, 2007  (9 PP), C. cattienensis Geissler, Nazarov, Orlov, Böhme, Phung, Nguyen & Ziegler, 2009  (6–8 PP), C. chauquangensis Hoang, Orlov, Ananjeva, Johns, Hoang & Dau, 2007  (6 or 7 PP), C. condorensis Smith 1921 a  (4–7 PP), C. cryptus Heidrich, Rösler, Vu, Böhme & Ziegler 2007  (9–11 PP), C. hontreensis Ngo, Grismer & Grismer 2008  (7 or 8 PP), C. huongsonensis Luu, Nguyen, Do & Ziegler 2011  (6 PP), C. huynhi Ngo & Bauer, 2008  (7–9 PP), C. intermedius Smith  (8–10 PP), C. irregularis Smith 1921 b  (5–7 PP), C. kingsadai Ziegler, Phung, Le & Nguyen 2013  (7–9 PP), C. martini Ngo 2011  (4 PP), C. phongnhakebangensis Ziegler, Rösler, Herrmann & Thanh 2002  (32–42 PP+FP), C. phuocbinhensis Nguyen, Le, Tran, Orlov, Lathrop, Macculloch, Le, Jin, Nguyen, Nguyen, Hoang, Che, Murphy & Zhang, 2013  (7 PP), C. phuquocensis Ngo, Grismer & Grismer, 2010  (7–9 PP), C. pseudoquadrivirgatus Rösler, Vu, Nguyen, Ngo & Ziegler, 2008  (5–9 PP), and C. roesleri Ziegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh & Schmitz 2010  (20–28 PP+FP), C. takouensis Ngo & Bauer 2008  (3 or 4 PP), C.taynguyenensis Nguyen, Le, Tran, Orlov, Lathrop, Macculloch, Le, Jin, Nguyen, Nguyen, Hoang, Che, Murphy & Zhang, 2013  (6 PP), C. thochuensis Ngo & Grismer 2012  (3–5 PP), C. yangbayensis Ngo & Chan, 2010  (6–8 PP) and C. ziegleri Nazarov, Orlov, Nguyen & Ho, 2008  (5–8 PP).

Cyrtodactylus thuongae  sp. nov. differs from C. badenensis Nguyen, Orlov & Darevsky 2006  , C. grismeri Ngo 2008  and C. nigriocularis Nguyen, Orlov & Darevsky 2006  by having enlarged femoral scales and lacking transversely enlarged subcaudals.

Cyrtodactylus thuongae  sp. nov. differs from C. cucphuongensis Ngo & Chan 2011  , by the absence of transversely enlarged subcaudals, having fewer enlarged femoral scales (2–4 vs. 14) and the presence of femoral and precloacal pores in males.

Cyrtodactylus thuongae  sp. nov. differs from C. eisenmanae Ngo, 2008  by having more dorsal tubercle rows (16–18 vs. 14), fewer ventral scale rows (29–44 vs. 44 or 45), the lack of transversely enlarged subcaudals, and the presence of femoral and precloacal pores in males.

Cyrtodactylus thuongae  sp. nov. differs from C. paradoxus ( Darevsky & Szczerbak 1997)  by the presence of femoral pores in males, the absence of distinctly enlarged subcaudal scales and a different body pattern (blotched vs. banded).

With respect to the remaining Cyrtodactylus  (except for zoogeographically distant species from the Papua- Australian region, the Solomon Islands, India, Sri Lanka, the Nicobar islands and Nepal), Cyrtodactylus thuongae  sp. nov. has no or one precloacal pore (PP), separated from a series of 0–3 femoral pores (FP) on each side in males (0 or 1 + 0–3 PP+FP) and thus differs from the following species that have more precloacal and femoral pores: C. agusanensis ( Taylor 1915)  (7–11 + 3–14), C. astrum Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012  (31–38 PP and FP in a continuous series), C. auribalteatus Sumontha, Panitvong & Deein, 2010  (6 + 4–5), C. australotitiwangsaensis Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012  (39–45), C. ayeyarwadyensis Bauer 2003  (10–28), C. batucolus Grismer, Chan, Grismer, Wood & Belabut 2008  (43–46), C. bintangrendah Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012  (41–46), C. bintangtinggi Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012  (38–41), C. brevipalmatus (Smith)  (7–10 + 6–7), C. chanhomeae Bauer, Sumontha & Pauwels 2003  (32–34), C. consobrinoides ( Annandale 1905)  (26), C. deveti ( Brongersma 1948)  (18–22), C. dumnuii Bauer, Kunya, Sumontha, Niyomwan, Pauwels, Chanhome & Kunya 2010  (5–6 + 6), C. feae ( Boulenger 1893)  (32), C. fumosus (Müller 1995)  , C. gansi Bauer 2003  (42–52), C. halmahericus Mertens 1929  (42–52), C. jarujini Ulber 1991  (42– 54), C. langkawiensis Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012  (30), C. lekaguli Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012  (30–36), C. lomyenensis Ngo & Pauwels 2010  (32–40), C. macrotuberculatus Grismer & Norhayati 2008  (28–40), C. phuketensis Sumontha, Pauwels, Kunya, Nitikul, Samphantamit & Grismer, 2012  (33– 36), C. pulchellus Gray 1827  (33–39), C. redimiculus King 1962  (5–8 + 8–9), C. seribuatensis Youmans & Grismer 2006  (40–44), C. surin  Chan-ard & Makchai, 2011 (34), C. tamaiensis Mahony 2009  (40), C. trilatofasciatus Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012  (41– 46), C. variegatus ( Blyth 1861)  (32) and C. wetariensis ( Dunn 1927)  (11–12 + 12–13).

Cyrtodactylus thuongae  sp. nov. has a series of 0–3 femoral pores on each side and no or one precloacal pore in males and thus differs from the following species which lack femoral pores and have higher precloacal pore counts: C. aequalis Bauer 2003  (9 PP), C. annandalei Bauer 2003  (11–12), C. annulatus ( Taylor 1915)  (6), C. baluensis ( Mocquard 1890)  (9–11), C. chrysopylos Bauer 2003  , (10), C. consobrinus ( Peters 1871)  (9–11), C. durio Grismer, Anuar, Quah, Muin, Onn, Grismer & Ahmad 2010  (12), C. elok Dring 1979  (7 or 8), C. erythrops Bauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome, & Kunya, 2009  (9), C. ingeri Hikida 1990  (8), C. interdigitalis Ulber 1993  (14), C. khasiensis ( Jerdon 1870)  (12–14), C. majulah Grismer, Wood & Lim, 2012  (7–11), C. mandalayensis Mahony 2009  (5), C. mimikanus ( Boulenger 1914)  (7–17), C. nuaulu Oliver, Edgar, Mumpuni, Iskandar & Lilley 2009  (6), C. payacola Johnson, Quah, Anuar, Muin, Wood, Grismer, Greer, Chan, Ahmad, Bauer & Grismer, 2012  (11 or 12), C. pantiensis Grismer, Chan, Grismer, Wood & Belabut 2008  (8 or 9), C. peguensis ( Boulenger 1893)  (7–9), C. russelli Bauer 2003  (15), C. slowinskii Bauer 2002  (9–11), C. spinosus Linkem, McGuire, Hayden, Setiadi, Bickford & Brown 2008  (12–13), C. sworderi ( Smith 1925)  (8 or 9), C. teyniei David, Nguyen, Schneider & Ziegler 2011  (14 in the female holotype), C. tigroides Bauer, Sumontha & Pauwels 2003  (7 or 9) C. wakeorum Bauer 2003  (12) and C. wayakonei Nguyen, Kingsada, Rösler, Auer & Ziegler 2010  (6– 8).

Cyrtodactylus thuongae  sp. nov. differs from the following Cyrtodactylus  species by the absence of precloacal and femoral pores in females: C. batik Iskandar, Rachmansah & Umilaela 2011  , C. darmandvillei ( Weber 1890)  , C. gordongekkoi ( Das 1994)  , C. jarakensis Grismer, Chan, Grismer, Wood & Belabut 2008  , C. laevigatus ( Darevsky 1964)  , C. thirakhupti Pauwels, Bauer, Sumontha & Chanhome 2004  and C. wallacei Hayden, Brown, Gillespie, Setiadi, Linkem, Iskandar, Umilaela, Bickford, Riyanto, Mumpuni & McGuire 2008  .

Cyrtodactylus thuongae  sp. nov. has 29–44 ventral scale rows at midbody and thus differs from C. agamensis ( Rösler & Glaw 2008)  (67), C. gubaot Welton, Siler, Linkem, Diesmos & Brown, 2010  (54–67), C. lateralis (Werner 1986)  (60–64), C. malayanus ( De Rooij 1915)  (58–62), C. mamanwa Welton, Siler, Linkem, Diesmos & Brown, 2010  (57–70), C. matsuii Hikida 1990  (51), C. sanook Pauwels, Sumontha, Latinne & Grismer, 2013  (27– 28), and C. s umuroi Welton, Siler, Linkem, Diesmos & Brown 2010 (53–58).

Cyrtodactylus thuongae  sp. nov. has 16–18 dorsal longitudinal tubercle rows and thus differs from C. brevidactylus Bauer 2002  (27), C. buchardi David, Teynié & Ohler 2005  (25), C. oldhami ( Theobald 1876)  (19– 24) which lack enlarged femoral scales, C. redimiculus King 1962  (14–16), which lacks a ventro-lateral fold, C. sumonthai Bauer, Pauwels & Chanhome 2002  (12) and C. quadrivirgatus Taylor 1962  , which lack femoral pores.

Cyrtodactylus thuongae  sp. nov. differs from C. pageli Schneider, Nguyen, Schmitz, Kingsada, Auer & Ziegler 2011  by lacking precloacal pores in females (vs. 5 or 6).

Cyrtodactylus thuongae  sp. nov. differs from C. aurensis Grismer 2005  , C. cavernicolus Inger & King 1961  , C. jambangan Welton, Siler, Diesmos & Brown 2010  , C. jellesmae (Boulenger 1887)  , C. marmoratus Gray 1831  , C. papilionoides Ulber & Grossmann 1991  , C. philippinicus ( Steindachner 1867)  , C. pubisulcus Inger 1958  C. rubidus ( Blyth 1861)  , C. semenanjungensis Grismer & Leong 2005  C. stresemanni Rösler & Glaw 2008  , C. tautbatorum Welton, Siler, Diesmos & Brown 2009  , C. tiomanensis Das & Lim 2000  , by the absence of a precloacal groove.

Cyrtodactylus thuongae  sp. nov. differs from C. yoshii Hikida 1990  and C. zhaoermii Shi & Zhao 2010  by the presence of femoral pores in males. The new species further differs from C. zhaoermii  by having fewer enlarged femoral scales (2–4 vs. 12–14).

Cyrtodactylus thuongae  sp. n. differs from C. leegrismeri Chan & Norhayati 2010  by the absence of enlarged subcaudals, having fewer precloacal pores in males (absent or one vs. six) and the presence of femoral pores in males.

Cyrtodactylus thuongae  sp. n. differs from C. angularis  ( Smith 1921 b) by the presence of enlarged femoral scales and by having fewer precloacal pores in males (absent or one vs. six).

Distribution. Cyrtodactylus thuongae  is currently known only from the type locality in Tay Ninh Province, Vietnam (see Fig. 5View FIGURE 5).

Etymology. The species name is dedicated to the wife of the first author, Dr. Thuong Thi Lien Nguyen from University of Food Industry, Ho Chi Minh City, Vietnam, for support of his field research in Vietnam.

Natural history notes. Cyrtodactylus thuongae  inhabits the same habitat with two other co-occurring cave dwelling species, C. badenensis Nguyen, Orlov & Darevsky, 2006  and C. nigriocularis Nguyen, Orlov & Darevsky, 2006  ( Fig. 2View FIGURE 2). Specimens were collected at night (ca. 22–24:00) on the rocky wall in a small cave, approximately 0.5–2.5 m above the ground. Most of the specimens were found inside the cave (ca. 2–5 m from the entrance) and only one was collected outside (ca. 1.5 m away from the entrance). The surrounding habitat was mixed forest of hardwood and vine, dominated by trees of the families Moraceae  , Fabaceae  , and Dipterocarpaceae  . C. badenensis  and C. nigriocularis  were also found on rocky cliffs of the cave, usually at heights below 1.5 m above the ground.

VNMN

Vietnam National Museum of Nature

ZFMK

Zoologisches Forschungsmuseum Alexander Koenig

Kingdom

Animalia

Phylum

Chordata

Class

Reptilia

Order

Squamata

Family

Gekkonidae

Genus

Cyrtodactylus

Loc

Cyrtodactylus thuongae

Phung, Trung My, Schingen, Mona Van, Ziegler, Thomas & Nguyen, Truong Quang 2014

2014
Loc

C. kingsadai

Ziegler, Phung, Le & Nguyen 2013

2013
Loc

C. phuocbinhensis

Nguyen, Le, Tran, Orlov, Lathrop, Macculloch, Le, Jin, Nguyen, Nguyen, Hoang, Che, Murphy & Zhang 2013

2013
Loc

C.taynguyenensis

Nguyen, Le, Tran, Orlov, Lathrop, Macculloch, Le, Jin, Nguyen, Nguyen, Hoang, Che, Murphy & Zhang 2013

2013
Loc

C. sanook

Pauwels, Sumontha, Latinne & Grismer 2013

2013
Loc

C. bidoupimontis

Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler 2012

2012
Loc

C. bugiamapensis

Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler 2012

2012
Loc

C. thochuensis

Ngo & Grismer 2012

2012
Loc

C. astrum

Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012

2012
Loc

C. australotitiwangsaensis

Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012

2012
Loc

C. bintangrendah

Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012

2012
Loc

C. bintangtinggi

Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012

2012
Loc

C. langkawiensis

Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012

2012
Loc

C. lekaguli

Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012

2012
Loc

C. phuketensis

Sumontha, Pauwels, Kunya, Nitikul, Samphantamit & Grismer 2012

2012
Loc

C. trilatofasciatus

Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012

2012
Loc

C. majulah

Grismer, Wood & Lim 2012

2012
Loc

C. payacola

Johnson, Quah, Anuar, Muin, Wood, Grismer, Greer, Chan, Ahmad, Bauer & Grismer 2012

2012
Loc

C. huongsonensis

Luu, Nguyen, Do & Ziegler 2011

2011
Loc

C. martini

Ngo 2011

2011
Loc

C. cucphuongensis

Ngo & Chan 2011

2011
Loc

C. teyniei

David, Nguyen, Schneider & Ziegler 2011

2011
Loc

C. batik

Iskandar, Rachmansah & Umilaela 2011

2011
Loc

C. pageli

Schneider, Nguyen, Schmitz, Kingsada, Auer & Ziegler 2011

2011
Loc

C. bichnganae

Ngo & Grismer 2010

2010
Loc

C. phuquocensis

Ngo, Grismer & Grismer 2010

2010
Loc

C. roesleri

Ziegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh & Schmitz 2010

2010
Loc

C. yangbayensis

Ngo & Chan 2010

2010
Loc

C. auribalteatus

Sumontha, Panitvong & Deein 2010

2010
Loc

C. dumnuii

Bauer, Kunya, Sumontha, Niyomwan, Pauwels, Chanhome & Kunya 2010

2010
Loc

C. lomyenensis

Ngo & Pauwels 2010

2010
Loc

C. durio

Grismer, Anuar, Quah, Muin, Onn, Grismer & Ahmad 2010

2010
Loc

C. wayakonei Nguyen, Kingsada, Rösler, Auer & Ziegler 2010

Nguyen, Kingsada, Rosler, Auer & Ziegler 2010

2010
Loc

C. gubaot

Welton, Siler, Linkem, Diesmos & Brown 2010

2010
Loc

C. mamanwa

Welton, Siler, Linkem, Diesmos & Brown 2010

2010
Loc

C. jambangan

Welton, Siler, Diesmos & Brown 2010

2010
Loc

C. zhaoermii

Shi & Zhao 2010

2010
Loc

C. leegrismeri

Chan & Norhayati 2010

2010
Loc

C. cattienensis Geissler, Nazarov, Orlov, Böhme, Phung, Nguyen & Ziegler, 2009

Geissler, Nazarov, Orlov, Bohme, Phung, Nguyen & Ziegler 2009

2009
Loc

C. tamaiensis

Mahony 2009

2009
Loc

C. erythrops

Bauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome 2009

2009
Loc

C. mandalayensis

Mahony 2009

2009
Loc

C. nuaulu

Oliver, Edgar, Mumpuni, Iskandar & Lilley 2009

2009
Loc

C. tautbatorum

Welton, Siler, Diesmos & Brown 2009

2009
Loc

C. hontreensis

Ngo, Grismer & Grismer 2008

2008
Loc

C. huynhi

Ngo & Bauer 2008

2008
Loc

C. pseudoquadrivirgatus Rösler, Vu, Nguyen, Ngo & Ziegler, 2008

Rosler, Vu, Nguyen, Ngo & Ziegler 2008

2008
Loc

C. takouensis

Ngo & Bauer 2008

2008
Loc

C. ziegleri

Nazarov, Orlov, Nguyen & Ho 2008

2008
Loc

C. grismeri

Ngo 2008

2008
Loc

C. eisenmanae

Ngo 2008

2008
Loc

C. batucolus

Grismer, Chan, Grismer, Wood & Belabut 2008

2008
Loc

C. macrotuberculatus

Grismer & Norhayati 2008

2008
Loc

C. pantiensis

Grismer, Chan, Grismer, Wood & Belabut 2008

2008
Loc

C. spinosus

Linkem, McGuire, Hayden, Setiadi, Bickford & Brown 2008

2008
Loc

C. jarakensis

Grismer, Chan, Grismer, Wood & Belabut 2008

2008
Loc

C. wallacei

Hayden, Brown, Gillespie, Setiadi, Linkem, Iskandar, Umilaela, Bickford, Riyanto, Mumpuni & McGuire 2008

2008
Loc

C. agamensis ( Rösler & Glaw 2008 )

Rosler & Glaw 2008

2008
Loc

C. stresemanni Rösler & Glaw 2008

Rosler & Glaw 2008

2008
Loc

C. caovansungi

Orlov, Nguyen, Nazarov, Ananjeva & Nguyen 2007

2007
Loc

C. chauquangensis

Hoang, Orlov, Ananjeva, Johns, Hoang & Dau 2007

2007
Loc

C. cryptus Heidrich, Rösler, Vu, Böhme & Ziegler 2007

Heidrich, Rosler, Vu, Bohme & Ziegler 2007

2007
Loc

C. badenensis

Nguyen, Orlov & Darevsky 2006

2006
Loc

C. nigriocularis

Nguyen, Orlov & Darevsky 2006

2006
Loc

C. seribuatensis

Youmans & Grismer 2006

2006
Loc

C. buchardi David, Teynié & Ohler 2005

David, Teynie & Ohler 2005

2005
Loc

C. aurensis

Grismer 2005

2005
Loc

C. semenanjungensis

Grismer & Leong 2005

2005
Loc

C. thirakhupti

Pauwels, Bauer, Sumontha & Chanhome 2004

2004
Loc

C. ayeyarwadyensis

Bauer 2003

2003
Loc

C. chanhomeae

Bauer, Sumontha & Pauwels 2003

2003
Loc

C. gansi

Bauer 2003

2003
Loc

C. aequalis

Bauer 2003

2003
Loc

C. annandalei

Bauer 2003

2003
Loc

C. chrysopylos

Bauer 2003

2003
Loc

C. russelli

Bauer 2003

2003
Loc

C. tigroides

Bauer, Sumontha & Pauwels 2003

2003
Loc

C. wakeorum

Bauer 2003

2003
Loc

C. phongnhakebangensis Ziegler, Rösler, Herrmann & Thanh 2002

Ziegler, Rosler, Herrmann & Thanh 2002

2002
Loc

C. slowinskii

Bauer 2002

2002
Loc

C. brevidactylus

Bauer 2002

2002
Loc

C. sumonthai

Bauer, Pauwels & Chanhome 2002

2002
Loc

C. tiomanensis

Das & Lim 2000

2000
Loc

C. paradoxus (

Darevsky & Szczerbak 1997

1997
Loc

C. fumosus (Müller 1995)

Muller 1995

1995
Loc

C. gordongekkoi (

Das 1994

1994
Loc

C. interdigitalis

Ulber 1993

1993
Loc

C. jarujini

Ulber 1991

1991
Loc

C. papilionoides

Ulber & Grossmann 1991

1991
Loc

C. ingeri

Hikida 1990

1990
Loc

C. matsuii

Hikida 1990

1990
Loc

C. yoshii

Hikida 1990

1990
Loc

C. lateralis

Werner 1986

1986
Loc

C. elok

Dring 1979

1979
Loc

C. laevigatus (

Darevsky 1964

1964
Loc

C. redimiculus

King 1962

1962
Loc

C. redimiculus

King 1962

1962
Loc

C. quadrivirgatus

Taylor 1962

1962
Loc

C. cavernicolus

Inger & King 1961

1961
Loc

C. pubisulcus

Inger 1958

1958
Loc

C. deveti (

Brongersma 1948

1948
Loc

C. halmahericus

Mertens 1929

1929
Loc

C. wetariensis (

Dunn 1927

1927
Loc

C. sworderi (

Smith 1925

1925
Loc

C. condorensis

Smith 1921

1921
Loc

C. irregularis

Smith 1921

1921
Loc

C. agusanensis (

Taylor 1915

1915
Loc

C. annulatus (

Taylor 1915

1915
Loc

C. malayanus (

De Rooij 1915

1915
Loc

C. mimikanus (

Boulenger 1914

1914
Loc

C. consobrinoides (

Annandale 1905

1905
Loc

C. feae (

Boulenger 1893

1893
Loc

C. peguensis (

Boulenger 1893

1893
Loc

C. baluensis (

Mocquard 1890

1890
Loc

C. darmandvillei (

Weber 1890

1890
Loc

C. jellesmae

Boulenger 1887

1887
Loc

C. oldhami (

Theobald 1876

1876
Loc

C. consobrinus (

Peters 1871

1871
Loc

C. khasiensis (

Jerdon 1870

1870
Loc

C. philippinicus (

Steindachner 1867

1867
Loc

C. variegatus (

Blyth 1861

1861
Loc

C. rubidus (

Blyth 1861

1861
Loc

C. marmoratus

Gray 1831

1831
Loc

C. pulchellus

Gray 1827

1827