Cyrtodactylus huongsonensis , Luu, Vinh Quang, Nguyen, Truong Quang, Do, Huy Quang & Ziegler, Thomas, 2011

Luu, Vinh Quang, Nguyen, Truong Quang, Do, Huy Quang & Ziegler, Thomas, 2011, A new Cyrtodactylus (Squamata: Gekkonidae) from Huong Son limestone forest, Hanoi, northern Vietnam, Zootaxa 3129, pp. 39-50: 40-48

publication ID

10.5281/zenodo.201564

publication LSID

lsid:zoobank.org:pub:B141FA6A-9430-48FA-A85E-123D64845E81

persistent identifier

http://treatment.plazi.org/id/C220867F-FFC3-0246-B9BA-05A23CFEB57A

treatment provided by

Plazi

scientific name

Cyrtodactylus huongsonensis
status

sp. nov.

Cyrtodactylus huongsonensis  sp. nov.

Holotype. Adult male ( IEBR A.2011.3) collected on 13 June 2011 by Vinh Quang Luu, Hieu Van Pham, and Nghia Van Ha in Huong Son limestone forest (25 o 35 ’N, 105 o 45 ’E), My Duc District, Hanoi, northern Vietnam at an elevation of ca. 120 m a.s.l. ( Figs. 1–4View FIGURE 1View FIGURE 2View FIGURE 3View FIGURE 4. A).

Paratype. Adult female ( ZFMKAbout ZFMK 92293) the same collection data as for the holotype ( Fig. 4 View Figure ).

Diagnosis. A medium-sized Cyrtodactylus  with a maximum SVL of 89.8 mm, distinguished from all congeners by the combination of the following characters: 1) dorsal pattern consisting of a dark nuchal loop, neck band and five in part irregularly shaped transverse body bands between limbs; 2) two enlarged lateral chinshields in contact with first postmental pair; 3) dorsal tubercles present on occiput, body, forearms, hind limbs and tail base; 4) 14–16 irregular dorsal tubercle rows; 5) ventrals in 41–48 longitudinal rows at midbody; 6) lateral skin folds present, lacking tubercles; 7) six precloacal pores plus in total 17 femoral pores in males separated by 8–12 poreless scales; 8) enlarged femoral scales present; 9) enlarged precloacal scales present; 10) precloacal groove absent; 11) 3 postcloacal spurs in males; 12) subcaudal scales transversally enlarged.

Description of holotype. Size medium (SVL 89.8 mm, TaL 78.0 mm, regenerated), distance from posterior corner of eye to anterior margin of ear including ciliaria 8.4 mm, maximum horizontal ear diameter 1.9 mm; for further measurements see Table 1.

Rostral wider than high (RW 3.7 mm, RH 2.2 mm, RW/RH 1.2) with an inverse Y-shaped median suture; supralabials 10 / 12; infralabials 10 / 11; nares bordered by rostral anteriorly, first supralabial laterally and four nasals posteriorly; supranasals separated from each other by two nasorostrals and a rectangular internasal; medial snout scales granular, those in contact with and nearby supralabials flattened and larger than medial scales; upper anterior ciliaries 3–4 times larger than posterior cilaries; head scales somewhat granular, smaller than median snout scales; anterior supraocular region, back of head and temporal region with rounded, conical tubercles, 3–6 times larger than surrounding scales; mental triangular, slightly wider than rostral; one pair of enlarged postmentals, longer than wide, in broad contact posteriorly; postmentals bordering mental anteriorly, first labials, one pair of enlarged gular scales, with four small gular scales in between them; dorsal scales somewhat granular, about as large as medial snout scales; dorsal tubercles round, conical, surrounded by 9–10 granular scales, tubercles forming approximately 14 irregular longitudinal rows at midbody; ventral scales smooth, medial scales 2–4 times larger than dorsal granules, 48 longitudinal rows at midbody; lateral folds present, without interspersed tubercles; upper arm lacking enlarged tubercles, lower arm with some in part indistinct conical tubercles; dorsal hind limb covered with scattered, smooth, flat to conical tubercles; femoral pores bearing scales distinctly enlarged, separated from precloacal, pore-bearing scales by 8 / 12 poreless femoral scales; 9 / 7 enlarged femoral scales, 10 / 7 femoral pores, six precloacal pores in an angular series; fingers and toes lacking distinct webbing; lamellae numbering 15 / 15 under first finger, 19 / 17 under fourth finger, 15 / 15 under first toe, and 20 / 20 under fourth toe; claws surrounded by a small scale on upper and a large scale on lower sides; precloacal region covered by a patch of approximately 29 enlarged scales below precloacal, pore-bearing scales; precloacal groove absent; 3 / 3 enlarged postcloacal tubercles on lateral surface of hemipenial swelling; dorsum of tail bearing distinct tubercles at base only, remainder of tail smooth, regenerated; subcaudals distinctly enlarged, flat, smooth.

Coloration in ethanol. Ground colouration light brownish-grey, with dark brownish-black dorsal pattern; dorsal head surface with irregular dark blotches, largest at occiput; dark canthal stripe, extending from nostril to eye; postocular streak only distinct on the right side, continuing to contact a somewhat irregularly shaped nuchal loop; postocular streak and nuchal loop bordered by thin light line; neck with a short dark transversal band; five more distinct dark transverse bands between limbs; dark transverse body bands somewhat irregularly shaped, with dark and light borders, fading towards sides; interspaces between dark dorsal bands with dark reticulation or blotches; dark tubercles in the nuchal loop and body bands, whereas light tubercles comprise the light lines and interspaces; flanks with dark blotches; upper surfaces of limbs with dark stripes and reticulations; dorsal surface of regenerated tail grey; gular region yellowish-cream; venter cream and lower tail surface light grey.

For coloration in life see Fig. 1 View Figure . The light lines bordering the nuchal loop and dark dorsal bands are yellow in life, as are the light dorsal tubercles; ground colouration in life is light greyish-brown.

Variation of paratype. For the variation in colouration of the female paratype see Fig. 4 View Figure and for meristics see Table 1. The internasal is very small and located between the anterior edges of the nasorostrals. The tail is original and has distinctly broadened subcaudals. The female paratype is somewhat smaller, but due to egg development we classify it as an adult, which is also indicated by the tail colouration. The dorsal surface of the original tail has 11 dark and 11 light bands, of which the last four dark bands are only faintly discernible. The disappearing light tail tip indicates the transition from juvenile to adult stage (see Ziegler et al. 2010). Concerning the colour pattern, the postocular streaks are well developed on both sides of the female paratype. Here, also the lateral dark blotches are more distinct than it is the case in the male holotype. The dark blotches on the occiput are fused to a black v-shaped transverse line. The underside of the tail is dark grey, with the tip being light. With respect to sexual dimorphism, the female lacks hemipenial swellings at the tail base, the precloacal and femoral pores are only faintly discernible, and the postcloacal tubercles are less pronounced and fewer in number (2 / 2 versus 3 / 3 in the male holotype).

Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Grismer et al. 2008, Rösler & Glaw 2008, Bauer et al. 2009, 2010, Ngo & Grismer 2010, Ngo & Pauwels 2010, Sumontha et al. 2010, Ziegler et al. 2010, David et al. 2011, Iskandar et al. 2011, Ngo 2011, Schneider et al. 2011).

Cyrtodactylus huongsonensis  sp. nov. differs from its Vietnamese congeners by the following characters (for details see Table 2): Cyrtodactylus huongsonensis  sp. nov. has enlarged subcaudal scales and thus differs from the following species, which lack enlarged subcaudals: C. cattienensis Geissler, Nazarov, Orlov, Böhme, Phung, Nguyen & Ziegler, 2009  , C. cryptus Heidrich, Rösler, Vu, Böhme & Ziegler, 2007  , C. huynhi Ngo & Bauer, 2008  , C. irregularis ( Smith, 1921)  , C. pseudoquadrivirgatus Rösler, Vu, Nguyen, Ngo & Ziegler, 2008  , and C. ziegleri Nazarov, Orlov, Nguyen & Ho, 2008  . Cyrtodactylus huongsonensis  sp. nov. has enlarged femoral scales, which are lacking in the following species: C. badenensis Nguyen, Orlov & Darevsky, 2006  , C. chauquangensis Hoang, Orlov, Ananjeva, Johns, Hoang & Dau, 2007  , C. grismeri Ngo, 2008  , and C. nigriocularis Nguyen, Orlov & Darevsky, 2006  . Cyrtodactylus huongsonensis  sp. nov. has femoral pores, which are absent in the following species: C. eisenmanae Ngo, 2008  , C. hontreensis Ngo, Grismer & Grismer, 2008  , C. intermedius (Smith, 1917)  , C. martini Ngo, 2011  , C. paradoxus ( Darevsky & Szczerbak, 1997)  , and C. phuquocensis Ngo, Grismer & Grismer, 2010  . Cyrtodactylus huongsonensis  sp. nov. has 15–17 femoral pores in total and thus differs from C. caovansungi Orlov, Nguyen, Nazarov, Ananjeva & Nguyen, 2007  (6), C. takouensis Ngo & Bauer, 2008  (0–2), and C. yangbayensis Ngo & Chan, 2010  (0–2). Cyrtodactylus huongsonensis  sp. nov. has precloacal pores separated from femoral pores, which are a contiguous series in C. phongnhakebangensis Ziegler, Rösler, Herrmann & Vu, 2003  , and C. roesleri Ziegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh & Schmitz, 2010  . Cyrtodactylus huongsonensis  sp. nov. has three postcloacal tubercles in males and 41–48 ventrals and thus differs from C. bichnganae Ngo, 2010  , that has only two postcloacal tubercles in males and 30–31 ventrals. Cyrtodactylus huongsonensis  sp. nov. lacks tubercles on the lateral skin fold and on the dorsal tail surface and thus differs from C. condorensis ( Smith, 1921)  ; in addition, the latter species only has precloacal pores (4–7) and a blotched dorsal pattern.

With respect to the remaining Cyrtodactylus  , Cyrtodactylus huongsonensis  sp. nov. has transversely enlarged subcaudals and thus differs from the following species which lack enlarged subcaudals: C. adleri Das, 1997  , C. aequalis Bauer, 2003  , C. agusanensis (Taylor, 1915)  , C. annulatus (Taylor, 1915)  , C. batucolus Grismer, Chan, Grismer, Wood & Belabut, 2008  , C. biordinis Brown & McCoy, 1980  , C. brevidactylus Bauer, 2002  , C. buchardi David, Teynié & Ohler, 2004  , C. cavernicolus Inger & King, 1961  , C. derongo Brown & Parker, 1973  , C. fumosus (Müller, 1895)  , C. gansi Bauer, 2003  , C. halmahericus Mertens, 1929  , C. irianjayaensis Rösler, 2000  , C. jambangan Welton, Siler, Diesmos & Brown  , C. jellesmae (Boulenger, 1897)  , C. lateralis (Werner, 1896)  , C. loriae (Boulenger, 1898)  , C. louisiadensis ( De Vis, 1892)  , C. malayanus ( De Rooij, 1915)  , C. mandalayensis Mahony, 2009  , C. marmoratus Gray, 1831  , C. matsuii Hikida, 1990  , C. murua Kraus & Allison, 2006  , C. novaeguineae (Schlegel, 1837)  , C. nuaulu Oliver, Edgar, Mumpuni, Iskandar & Lilley, 2009  , C. pantiensis Grismer, Chan, Grismer, Wood & Belabut, 2008  , C. papuensis (Brongersma, 1934)  , C. philippinicus (Steindachner, 1867)  , C. pubisulcus Inger, 1957  , C. quadrivirgatus Taylor, 1962  , C. sadleiri Wells & Wellington, 1985  , C. semenanjungensis Grismer & Leong, 2005  , C. seribuatensis Youmans & Grismer, 2006  , C. sermowaiensis  (de Rooij, 1915), C. serratus Kraus, 2007  , C. stresemanni Rösler & Glaw, 2008  , C. sworderi (Smith, 1925)  , C. tautbatorum Welton, Siler, Diesmos & Brown, 2009  , C. tiomanensis Das & Lim, 2000  , C. tuberculatus (Lucas & Frost, 1900)  , C. wakeorum Bauer 2003  , C. wetariensis (Dunn, 1927)  , C. yoshii Hikida, 1990  , C. zhaoermii Shi & Zhao, 2010  , and C. zugi Oliver, Tjarhan, Mumpuni, Krey & Richards, 2008  .

Cyrtodactylus huongsonensis  sp. nov. has femoral pores and thus differs from the following species which lack femoral pores: C. angularis ( Smith, 1921)  , C. aurensis Grismer, 2005  , C. ayeyarwadyensis Bauer, 2003  , C. chanhomeae Bauer, Sumontha & Pauwels, 2003  , C. chrysopylos Bauer, 2003  , C. consobrinoides (Annandale, 1905)  , C. cracens Batuwita & Bahir, 2005  , C. deveti (Brongersma, 1948)  , C. edwardtaylori Batuwita & Bahir, 2005  , C. elok Dring, 1979  , C. feae (Boulenger, 1893)  , C. fraenatus (Günther, 1864)  , C. ingeri Hikida, 1990  , C. jarujini Ulber, 1993  , C. khasiensis (Jerdon, 1870)  , C. malcolmsmithi (Constable, 1949)  , C. oldhami (Theobald, 1876)  , C. pageli Schneider, Nguyen, Schmitz, Kingsada, Auer & Ziegler, 2011  , C. papilionoides Ulber & Grossmann, 1991  , C. peguensis (Boulenger, 1893)  , C. ramboda Batuwita & Bahir, 2005  , C. rubidus (Blyth, 1860)  , C. soba Batuwita & Bahir, 2005  , C. subsolanus Batuwita & Bahir, 2005  , C. sumonthai Bauer, Pauwels & Chanhome, 2002  , and C. variegatus (Blyth, 1859)  .

Cyrtodactylus huongsonensis  sp. nov. has six precloacal pores plus in total 17 femoral pores in males, which are separated by poreless scales and thus differs from the following species which have a contiguous series of precloacal-femoral pores: C. epiroticus Kraus, 2008  (60–82), C. klugei Kraus, 2008  (66–76), C. lomyenensis Ngo & Pauwels, 2010  (32–40), C. robustus Kraus, 2008  (75–85), C. salomonensis Rösler, Richards & Günther, 2007  (71– 72), C. tamaiensis Mahony, 2009  (40), and C. tripartitus Kraus, 2008  (64–78).

Cyrtodactylus huongsonensis  sp. nov. has six precloacal pores in males and 8 precloacal pores in females and thus differs from the following species which have distinctly higher precloacal pore counts: C. annandalei Bauer, 2003  (11–12), C. capreoloides Rösler, Richards & Günther, 2007  (13), C. durio Grismer et al., 2010  (12), C. interdigitalis Ulber, 1993  (14), C. russelli Bauer, 2003  (15), C. spinosus Linkem, McGuire, Hayden, Setiadi, Bickford & Brown  (12–13), and C. teyniei David, Nguyen, Schneider & Ziegler, 2011  (14 in the single known specimen, an adult female).

The following Cyrtodactylus  species differ from Cyrtodactylus huongsonensis  sp. nov. by the absence of precloacal and femoral pores in both sexes: C. batik Iskandar, Rachmansah & Umilaela, 2011  , C. darmandvillei (Weber, 1890)  , C. gordongekkoi (Das, 1993)  (see Biswas 2007), C. jarakensis Grismer, Chan, Grismer, Wood & Belabut, 2008  , C. laevigatus (Darevsky, 1964)  , C. thirakhupti Pauwels, Bauer, Sumontha & Chanhome 2004  , and C. wallacei Hayden, Brown, Gillespie, Setiadi, Linkem, Iskandar, Umilaela, Bickford, Riyanto, Mumpuni & McGuire, 2008  .

Cyrtodactylus huongsonensis  sp. nov. has 41–48 ventral scales at midbody and thus differs from C. agamensis (Bleeker, 1860)  (67), C. consobrinus (Peters, 1871)  (65–70), C. erythrops Bauer et al., 2008  (28), C. gubernatoris (Annandale, 1913)  (30), C. leegrismeri Chan & Norhayati, 2010  (27–35), C. macrotuberculatus Grismer & Norhayati, 2008  (19–22), C. pulchellus Gray, 1828  (33–35), C. slowinskii Bauer, 2002  (27–32), C. tigroides Bauer, Sumontha & Pauwels, 2003  (34), and C. wayakonei Nguyen, Kingsada, Rösler, Auer & Ziegler, 2010  (31–35).

Cyrtodactylus huongsonensis  sp. nov. has dorsal tail surface without tubercles and thus differs from C. aaroni Günther & Rösler, 2003  , C. brevipalmatus (Smith, 1923)  , C. redimiculus King, 1962  , C. baluensis (Mocquard, 1890)  , and C. mimikanus (Boulenger, 1914)  .

Cyrtodactylus huongsonensis  sp. nov. has 14–16 dorsal tubercle rows and thus differs from C. auribalteatus Sumontha, Panitvong & Deein, 2010  (22–24), and C. dumnuii Bauer, Kunya, Sumontha, Niyomwan, Pauwels, Chanhome & Kunya, 2010  (18–22).

In addition, Cyrtodactylus huongsonensis  sp. n. differs from the representatives of the subgenus Geckoella  by the presence of transversally enlarged subcaudals and by having precloacal pores in both sexes: C. (Geckoella) albofasciatus (Boulenger, 1885)  , C. (Geckoella) collegalensis (Beddome, 1870)  , C. (Geckoella) deccanensis (Günther, 1864)  , C. (Geckoella) jeyporensis (Beddome, 1877)  , C. (Geckoella) nebulosus (Beddome, 1870)  , C. (Geckoella) triedrus (Günther, 1864)  , and C. (Geckoella) yakhuna (Deraniyagala, 1945)  .

Distribution. Cyrtodactylus huongsonensis  sp. nov. is currently known only from the type locality in Vietnam ( Fig. 5 View Figure ).

Etymology. We name this species after its type locality, Huong Son limestone forest, which is a tourist site and not yet protected. Huong Son tourist site covers an area of a thousand hectares and includes a complex of mountains, rivers and streams, villages, pagodas, and grottoes surrounded by the Huong Tich Mountain Range, north of the Truong Son Range.

Ecological notes. The type specimens were found at night on karst outcrops in limestone forest, at an elevation of ca. 120 m a.s.l. The left testis of the male holotype measured 8.0 mm in length. The somewhat smaller female paratype contained further developed eggs of up to 3.4 mm diameter.

TABLE 1. Selected measurements and morphological characters of the male holotype and female paratype of Cyrtodactylus huongsonensis sp. nov.; measurements in mm, * = regenerated tail.

  IEBR A.2011.3 ZFMK 92293
SVL 89.8 73.4
TaL 78.0* 90.5
HH 10.1 9.0
HL 24.7 22.2
HW 17.2 15.1
OD 5.6 5.1
SE 10.2 9.9
SL 10/12 13/12
IL 10/11 12/12
N 4/4 4/4
IN 1 1
CIL 26/26 28/27
PM 2 2
DTR 14 16
GST 8-10 8-10
V 48 41
SLB 191 170
EFS 9/7 9/9
FP 10/7 8/7
PP 6 8
PAT 3 /3 2 /2
LD4 19/17 19/19
LT4 20/20 23/22

TABLE 1. Selected measurements and morphological characters of the male holotype and female paratype of Cyrtodactylus huongsonensis sp. nov.; measurements in mm, * = regenerated tail.

  IEBR A.2011.3 ZFMK 92293
SVL 89.8 73.4
TaL 78.0* 90.5
HH 10.1 9.0
HL 24.7 22.2
HW 17.2 15.1
OD 5.6 5.1
SE 10.2 9.9
SL 10/12 13/12
IL 10/11 12/12
N 4/4 4/4
IN 1 1
CIL 26/26 28/27
PM 2 2
DTR 14 16
GST 8-10 8-10
V 48 41
SLB 191 170
EFS 9/7 9/9
FP 10/7 8/7
PP 6 8
PAT 3 /3 2 /2
LD4 19/17 19/19
LT4 20/20 23/22
ZFMK

Zoologisches Forschungsmuseum Alexander Koenig

Kingdom

Animalia

Phylum

Chordata

Class

Reptilia

Order

Squamata

Family

Gekkonidae

Genus

Cyrtodactylus

Loc

Cyrtodactylus huongsonensis

Luu, Vinh Quang, Nguyen, Truong Quang, Do, Huy Quang & Ziegler, Thomas 2011

2011
Loc

C. martini

Ngo 2011

2011
Loc

C. pageli

Schneider, Nguyen, Schmitz, Kingsada, Auer & Ziegler 2011

2011
Loc

C. teyniei

David, Nguyen, Schneider & Ziegler 2011

2011
Loc

C. batik

Iskandar, Rachmansah & Umilaela 2011

2011
Loc

C. phuquocensis

Ngo, Grismer & Grismer 2010

2010
Loc

C. yangbayensis

Ngo & Chan 2010

2010
Loc

C. roesleri

Ziegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh & Schmitz 2010

2010
Loc

C. bichnganae

Ngo 2010

2010
Loc

C. zhaoermii

Shi & Zhao 2010

2010
Loc

C. lomyenensis

Ngo & Pauwels 2010

2010
Loc

C. durio

Grismer et al. 2010

2010
Loc

C. leegrismeri

Chan & Norhayati 2010

2010
Loc

C. wayakonei Nguyen, Kingsada, Rösler, Auer & Ziegler, 2010

Nguyen, Kingsada, Rosler, Auer & Ziegler 2010

2010
Loc

C. auribalteatus

Sumontha, Panitvong & Deein 2010

2010
Loc

C. dumnuii

Bauer, Kunya, Sumontha, Niyomwan, Pauwels, Chanhome & Kunya 2010

2010
Loc

C. cattienensis Geissler, Nazarov, Orlov, Böhme, Phung, Nguyen & Ziegler, 2009

Geissler, Nazarov, Orlov, Bohme, Phung, Nguyen & Ziegler 2009

2009
Loc

C. mandalayensis

Mahony 2009

2009
Loc

C. nuaulu

Oliver, Edgar, Mumpuni, Iskandar & Lilley 2009

2009
Loc

C. tautbatorum

Welton, Siler, Diesmos & Brown 2009

2009
Loc

C. tamaiensis

Mahony 2009

2009
Loc

C. huynhi

Ngo & Bauer 2008

2008
Loc

C. pseudoquadrivirgatus Rösler, Vu, Nguyen, Ngo & Ziegler, 2008

Rosler, Vu, Nguyen, Ngo & Ziegler 2008

2008
Loc

C. ziegleri

Nazarov, Orlov, Nguyen & Ho 2008

2008
Loc

C. grismeri

Ngo 2008

2008
Loc

C. eisenmanae

Ngo 2008

2008
Loc

C. hontreensis

Ngo, Grismer & Grismer 2008

2008
Loc

C. takouensis

Ngo & Bauer 2008

2008
Loc

C. batucolus

Grismer, Chan, Grismer, Wood & Belabut 2008

2008
Loc

C. pantiensis

Grismer, Chan, Grismer, Wood & Belabut 2008

2008
Loc

C. stresemanni Rösler & Glaw, 2008

Rosler & Glaw 2008

2008
Loc

C. zugi

Oliver, Tjarhan, Mumpuni, Krey & Richards 2008

2008
Loc

C. epiroticus

Kraus 2008

2008
Loc

C. klugei

Kraus 2008

2008
Loc

C. robustus

Kraus 2008

2008
Loc

C. tripartitus

Kraus 2008

2008
Loc

C. jarakensis

Grismer, Chan, Grismer, Wood & Belabut 2008

2008
Loc

C. wallacei

Hayden, Brown, Gillespie, Setiadi, Linkem, Iskandar, Umilaela, Bickford, Riyanto, Mumpuni & McGuire 2008

2008
Loc

C. erythrops

Bauer et al. 2008

2008
Loc

C. macrotuberculatus

Grismer & Norhayati 2008

2008
Loc

C. cryptus Heidrich, Rösler, Vu, Böhme & Ziegler, 2007

Heidrich, Rosler, Vu, Bohme & Ziegler 2007

2007
Loc

C. chauquangensis

Hoang, Orlov, Ananjeva, Johns, Hoang & Dau 2007

2007
Loc

C. caovansungi

Orlov, Nguyen, Nazarov, Ananjeva & Nguyen 2007

2007
Loc

C. serratus

Kraus 2007

2007
Loc

C. salomonensis Rösler, Richards & Günther, 2007

Rosler, Richards & Gunther 2007

2007
Loc

C. capreoloides Rösler, Richards & Günther, 2007

Rosler, Richards & Gunther 2007

2007
Loc

C. badenensis

Nguyen, Orlov & Darevsky 2006

2006
Loc

C. nigriocularis

Nguyen, Orlov & Darevsky 2006

2006
Loc

C. murua

Kraus & Allison 2006

2006
Loc

C. seribuatensis

Youmans & Grismer 2006

2006
Loc

C. semenanjungensis

Grismer & Leong 2005

2005
Loc

C. aurensis

Grismer 2005

2005
Loc

C. cracens

Batuwita & Bahir 2005

2005
Loc

C. edwardtaylori

Batuwita & Bahir 2005

2005
Loc

C. ramboda

Batuwita & Bahir 2005

2005
Loc

C. soba

Batuwita & Bahir 2005

2005
Loc

C. subsolanus

Batuwita & Bahir 2005

2005
Loc

C. buchardi David, Teynié & Ohler, 2004

David, Teynie & Ohler 2004

2004
Loc

C. thirakhupti

Pauwels, Bauer, Sumontha & Chanhome 2004

2004
Loc

C. phongnhakebangensis Ziegler, Rösler, Herrmann & Vu, 2003

Ziegler, Rosler, Herrmann & Vu 2003

2003
Loc

C. aequalis

Bauer 2003

2003
Loc

C. gansi

Bauer 2003

2003
Loc

C. wakeorum

Bauer 2003

2003
Loc

C. ayeyarwadyensis

Bauer 2003

2003
Loc

C. chanhomeae

Bauer, Sumontha & Pauwels 2003

2003
Loc

C. chrysopylos

Bauer 2003

2003
Loc

C. annandalei

Bauer 2003

2003
Loc

C. russelli

Bauer 2003

2003
Loc

C. tigroides

Bauer, Sumontha & Pauwels 2003

2003
Loc

C. aaroni Günther & Rösler, 2003

Gunther & Rosler 2003

2003
Loc

C. brevidactylus

Bauer 2002

2002
Loc

C. sumonthai

Bauer, Pauwels & Chanhome 2002

2002
Loc

C. slowinskii

Bauer 2002

2002
Loc

C. irianjayaensis Rösler, 2000

Rosler 2000

2000
Loc

C. tiomanensis

Das & Lim 2000

2000
Loc

C. paradoxus (

Darevsky & Szczerbak 1997

1997
Loc

C. adleri

Das 1997

1997
Loc

C. jarujini

Ulber 1993

1993
Loc

C. interdigitalis

Ulber 1993

1993
Loc

C. gordongekkoi

Das 1993

1993
Loc

C. papilionoides

Ulber & Grossmann 1991

1991
Loc

C. matsuii

Hikida 1990

1990
Loc

C. yoshii

Hikida 1990

1990
Loc

C. ingeri

Hikida 1990

1990
Loc

C. sadleiri

Wells & Wellington 1985

1985
Loc

C. biordinis

Brown & McCoy 1980

1980
Loc

C. elok

Dring 1979

1979
Loc

C. derongo

Brown & Parker 1973

1973
Loc

C. laevigatus

Darevsky 1964

1964
Loc

C. quadrivirgatus

Taylor 1962

1962
Loc

C. redimiculus

King 1962

1962
Loc

C. cavernicolus

Inger & King 1961

1961
Loc

C. pubisulcus

Inger 1957

1957
Loc

C. malcolmsmithi

Constable 1949

1949
Loc

C. deveti

Brongersma 1948

1948
Loc

C. (Geckoella) yakhuna

Deraniyagala 1945

1945
Loc

C. papuensis

Brongersma 1934

1934
Loc

C. halmahericus

Mertens 1929

1929
Loc

C. wetariensis

Dunn 1927

1927
Loc

C. sworderi

Smith 1925

1925
Loc

C. brevipalmatus

Smith 1923

1923
Loc

C. irregularis (

Smith 1921

1921
Loc

C. condorensis (

Smith 1921

1921
Loc

C. angularis (

Smith 1921

1921
Loc

C. intermedius

Smith 1917

1917
Loc

C. agusanensis

Taylor 1915

1915
Loc

C. annulatus

Taylor 1915

1915
Loc

C. malayanus (

De Rooij 1915

1915
Loc

C. mimikanus

Boulenger 1914

1914
Loc

C. gubernatoris

Annandale 1913

1913
Loc

C. consobrinoides

Annandale 1905

1905
Loc

C. tuberculatus

Lucas & Frost 1900

1900
Loc

C. loriae

Boulenger 1898

1898
Loc

C. jellesmae

Boulenger 1897

1897
Loc

C. lateralis

Werner 1896

1896
Loc

C. fumosus (Müller, 1895)

Muller 1895

1895
Loc

C. feae

Boulenger 1893

1893
Loc

C. peguensis

Boulenger 1893

1893
Loc

C. louisiadensis (

De Vis 1892

1892
Loc

C. darmandvillei

Weber 1890

1890
Loc

C. baluensis

Mocquard 1890

1890
Loc

C. (Geckoella) albofasciatus

Boulenger 1885

1885
Loc

C. (Geckoella) jeyporensis

Beddome 1877

1877
Loc

C. oldhami

Theobald 1876

1876
Loc

C. consobrinus

Peters 1871

1871
Loc

C. khasiensis

Jerdon 1870

1870
Loc

C. (Geckoella) collegalensis

Beddome 1870

1870
Loc

C. (Geckoella) nebulosus

Beddome 1870

1870
Loc

C. philippinicus

Steindachner 1867

1867
Loc

C. fraenatus (Günther, 1864)

Gunther 1864

1864
Loc

C. (Geckoella) deccanensis (Günther, 1864)

Gunther 1864

1864
Loc

C. (Geckoella) triedrus (Günther, 1864)

Gunther 1864

1864
Loc

C. rubidus

Blyth 1860

1860
Loc

C. agamensis

Bleeker 1860

1860
Loc

C. variegatus

Blyth 1859

1859
Loc

C. novaeguineae

Schlegel 1837

1837
Loc

C. marmoratus

Gray 1831

1831
Loc

C. pulchellus

Gray 1828

1828