Volovnik, Semyon V., Boldgiv, Bazartseren, Iderzorig, Badamnyambuu, Khobrakova, Larisa Ts., Kolov, Sergey V., Rudykh, Sergey G., Zabaluev, Ilya A. & Grebennikov, Vasily V., 2021, The first molecular phylogeny of the weevil subfamily Lixinae (Coleoptera Curculionidae) casts doubts on the monophyly of its tribes, Zootaxa 5026 (2), pp. 201-220: 216

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Aside: Cleonini   are not universally flightless

Regardless of whether the tribe is monophyletic or not, some Cleonini   species do actively fly. This is, however, a poorly documented and rarely observed phenomenon, which misled Arzanov & Grebennikov (2017) into the erroneous conclusion that all Cleonini   are flightless. Active flight has been reported for at least three economically important species: Asproparthenis punctiventris (Germar, 1823)   (e.g., Orlachiova 1956; Auersch 1961; Tóth et al. 2006), Bothynoderes affinis (Schrank, 1781) ( Kokovikhin 1966)   , and Temnorhinus brevirostris (Gyllenhal, 1834) ( Brunner 1959)   . Meregalli (2005) reported flight in Eurycleonus talamellii Meregalli, 2005   . One of us (BI) observed in Mongolia active flight of, likely, Conorhynchus conirostris Gebler, 1829   . Robert Stejskal (personal communication) observed in central Europe active flight of Bothynoderes affinis   , Asproparthenis punctiventris   , Cleonis pigra   , and Cyphocleonus dealbatus Gmelin, 1790   .