Graphicomassa korytnicensis ( Bałuk, 1995 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard M., 2021, The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group, Zootaxa 5025 (1), pp. 1-75 : 12-13

Graphicomassa korytnicensis ( Bałuk, 1995) nov. comb.

Figs 3G 1 –G 2 , H 1 –H 2 , I 1 –I 2 , J 1 –J 2 , K, L

Columbella curta Bell. —Hörnes 1852: 118 (pars), pl. 11, figs 4–6 [non Dujardin, 1837].

Columbella curta Dujardin — Hoernes & Auinger 1880 (pars): 93, pl. 7, figs 15–19 [non Dujardin, 1837].

Columbella (Columbella) curta Duj. — Boettger 1902: 16 [non Dujardin, 1837].

Columbella (Mitrella) curta Duj. — Boettger 1906: 14 [non Dujardin, 1837].

Columbella (Mitrella) an turgidula Brocc. —Friedberg 1928: 586, pl. 38, fig. 2.

Columbella an turgidula Brocc. — Friedberg 1938: 128.

Columbella (Alia) curta var. convexa Friedberg 1911 —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 182, pl. 45, fig. 3 [non Mitrella convexa ( Friedberg, 1911) ].

* Pyrene (Mitrella) korytnicensis sp. n. — Bałuk 1995: 236, pl. 31, figs 11–12, pl. 32, fig. 7.

Type material. Holotype: Institute of Geology of the University of Warsaw ( ZPALUW BkK-G619), SL: 16.0 mm, MD: 8.5 mm, Korytnica, Poland, illustrated in Bałuk (1995, pl. 31, fig. 12).

Illustrated material. NHMW1858 /0043/0016, SL: 21.4 mm, MD: 11.4 mm, Lăpugiu de Sus ( Romania), Figs 3J 1 –J 2 . NHMW1854 /0035/0106, SL: 23.4 mm, MD: 11.2 mm, Lăpugiu de Sus ( Romania), Fig. 3K . NHMW1854 / 0035/0167a, SL: 20.1 mm, MD: 10.4 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 15a–b), Figs 3I 1 –I 2 . NHMW1854 /0035/016c7, SL: 18.9 mm, MD: 10.6 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 17a–b), Figs 3G 1 –G 2 . NHMW1854 /0035/0167d, SL: 15.9 mm, MD: 7.9 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 18a–c), Figs 3H 1 –H 2 ; NHMW2020 /0153/0001, Lăpugiu de Sus ( Romania), Fig 3L.

Additional material. NHMW1846 /0037/0155a, SL: 15.9 mm, MD: 7.0 mm, Bad Vöslau ( Austria) , illustrated in Hörnes (1852, pl. 11, fig. 5). NHMW1846 /0037/0155b, SL: 13.6 mm, MD: 6.6 mm, Bad Vöslau ( Austria) , illustrated in Hörnes (1852, pl. 11, fig. 6). NHMW1854 /0035/0167b, SL: 20.4 mm, MD: 10.5 mm, Lăpugiu de Sus ( Romania) , illustrated in Hoernes & Auinger (1880, pl. 7, figs. 16a–b). NHMW1854 /0035/0167e, SL: 11.4 mm, MD: 5.5 mm, Lăpugiu de Sus ( Romania) , illustrated in Hoernes & Auinger (1880, pl. 7, figs. 19a–c). NHMW1869 /0001/0024, 12 spec., Bad Vöslau ( Austria) ; NHMW1868 /0001/0066, 5 spec., Baden-Sooss ( Austria) ; NHMW1872 /0030/0118, 1 spec., Baden ( Austria) ; NHMW1870 /0033/0041, 74 spec., Lăpugiu de Sus ( Romania) ; NHMW2020 /0153/0001, 6 spec., Lăpugiu de Sus ( Romania) ; NHMW 2020 /0172/0005 (ex B. Landau coll.), 2 spec., Lăpugiu de Sus ( Romania) .

Revised description. Shell large, robust, broad ovoid, with decollate apex and incised suture. Protoconch unknown. Spire moderately low, conical to cyrtoconoid, weakly gradate. Spire whorls flat-sided to weakly convex with narrow shoulder and periphery at abapical suture. Last whorl high, subcylindrical to weakly convex, rarely slightly pinched below shoulder. Base weakly to moderately constricted. Shell surface smooth except for few weak spiral cords on base.Aperture moderately narrow.Anal canal sharply incised, V-shaped, callused. Outer lip thickened with weakly developed internal labial callus pad in upper third and straight margin, denticulate with about 10–12 small denticles within. Siphonal canal short, wide, moderately deeply notched at tip. Columellar callus strongly thickened forming sharply delimited callus rim, slightly detached over base, abapically thickened with about seven indistinct denticles at outer columellar edge. Callus thins mid-aperture, thickening again in parietal area. Colour pattern of close-set, roughly spirally arranged, small subtrigonal whitish dots on brownish ground.

Shell measurements and ratios. SL: 15.9–23.4 mm, MD: 7.0– 11.4 mm, AA: 53–61°, SL/MD: 1.9, AL/AW: 3.5–3.6, LWH/AH: 1.3.

Discussion. This species has been mixed with Graphicomassa polonica by Hörnes (1852) and Hoernes & Auinger (1880). It differs from G. polonica in its stout outline, cyrtoconoid spire and shorter last whorl. In addition, the colour pattern of G. korytnicensis is delicate, comprising small dots, whereas G. polonica shows broad squarish flammulae. Nevertheless, subadult shells of Graphicomassa polonica can be difficult to distinguish from G. korytnicensis . Both species seem to have been ecologically separated. Graphicomassa polonica is found in inner neritic silty-sandy deposits whereas G. korytnicensis is found in offshore clays. The stout, thickened shell and cyrtoconoid spire make this species superficially similar to Columbella mayeri Hoernes, 1856 , but it is readily separated by the absence of the row of inner columellar denticles, diagnostic to the genus (see above). Moreover, the spire is even lower and more strongly cyrtoconoid in C. mayeri .

Palaeoenvironment. Middle neritic to outer neritic. The specimens from the Vienna Basin are found in clay of the Baden Fm., suggesting about 250 m water depth ( Hohenegger et al. 2008).

Distribution in Central Paratethys. Badenian (middle Miocene): Carpathian Foredeep: Korytnica ( Poland) ( Bałuk 1995); Zborów ( Ukraine) ( Friedberg 1938); Vienna Basin: Baden, Bad Vöslau, Baden-Sooss (own data); Făget Basin: Lăpugiu de Sus, Coşteiu de Sus ( Romania) ( Hoernes & Auinger 1880; Boettger 1906); Dacian Basin: Târnene ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width.