López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision

López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119

publication ID

https://doi.org/10.11646/zootaxa.5211.1.1

link to original document

https://doi.org/10.11646/zootaxa.5211.1.1

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Tomarus key    16
Euligyrus     17-18
Euligyrus ebenus   comb. nov.  18-19
Euligyrus similis   comb. nov.  19-21
Proculigyrus cicatricosus   comb. nov.  21-22
Ligyrus     22-24
Ligyrodes (Anagrylius)     24-25
Ligyrus (Anagrylius) cuniculus   comb. nov.  25-27
Ligyrus (Anagrylius) moroni   comb. nov.  28
Ligyrodes     29-30
Ligyrus (Ligyrodes) peruvianus     30
Ligyrus (Ligyrodes) relictus     31-32
Ligyrus (Ligyrodes) sallaei     32-35
Ligyrus     35-36
Ligyrus (Ligyrus) allonasutus   sp. nov.  37-38
Ligyrus (Ligyrus) burmeisteri     39-40
Ligyrus (Ligyrus) bidentulus     39
Ligyrus (Ligyrus) fossor     41-42
Ligyrus (Ligyrus) gibbosus     42-45
Ligyrus (Ligyrus) nasutus     46-47
Ligyrus (Ligyrus) neglectus     47
Ligyrus (Ligyrus) paranaensis   comb. nov.  48
Ligyrus (Ligyrus) peninsularis   comb. nov.  48-49
Ligyrus (Ligyrus) rubripes     49-50
Ligyrus (Ligyrus) ruginasus     50-51
Ligyrus (Ligyrus) villosus     51-52
Ligyrus (Ligyrus) spinipenis   comb. nov.  51
Tomarus     52-54
Tomarus amazonicus   new status and combination  55-56
Tomarus adoceteus     55
Tomarus bituberculatus     56-58
Tomarus discrepans     59-60
Tomarus colombianus     59
Tomarus gyas     60-62
Tomarus laevicollis     62-63
Tomarus laticaudus     63-64
Tomarus maimon     64-65
Tomarus maracaiboensis     65-66
Tomarus maternus     66-67
Tomarus pullus     67-68
Tomarus pilcopataensis     67
Tomarus pumilus     68-69
Tomarus roigjunenti     69-70
Tomarus rostratus     70
Tomarus selanderi     70-71
Tomarus subtropicus     71-72